Osmotic Range principle
Lessons from Bull Shark
Cross the water line.
River-Sea Movement

Core lesson
Real range comes from being able to change water without losing power.
Biological basis
Bull Sharks tolerate both salt and fresh water through osmoregulation and can move far into rivers as well as coastal marine habitats.
Best use cases
Where this lesson tends to be most useful in practice.
RangeAdaptationOsmoregulationBoundary crossingUnexpected paths