Dual Refuge principle
Lessons from Dusky Gopher Frog
Keep two homes.
Burrow-And-Pond Life Cycle

Core lesson
Some lives survive because they know which home to use in each season.
Biological basis
Dusky Gopher Frogs use upland burrows, often made by other animals, for shelter and migrate to temporary ponds for breeding.
Best use cases
Where this lesson tends to be most useful in practice.
RefugeLife cyclesSeasonal movementShelterAdapted timing