Two-World Belonging principle
Lessons from Tiger Salamander
Belong in both.
Burrow-to-Pool Life Cycle

Core lesson
Flexibility lets one life belong underground and in water.
Biological basis
Tiger Salamanders live much of the year underground or under cover and migrate to ponds or wetlands to breed, with aquatic larval stages.
Best use cases
Where this lesson tends to be most useful in practice.
FlexibilityLife cyclesTransitionBurrowingBreeding return